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Lacey Chabert as Eliza Thornberry

Tom Kane as Darwin Thornberry

Zion Acie as Tally

Rupert Everett as Sloan Blackburn

Taylor Swift as Bree Blackburn/Marianne Thornberry

Tim Curry as Nigel Thornberry

Tom Kenny as Donnie Thornberry

Danielle Harris as Debbie Thornberry

Jeff Bennett as Kovu Dominic Jr.

Cameron Boyce as Kovu Dominic III

Tara Strong as Mary Dominic